Roger L. Mann

Education

• Ph.D.  Marine Science. University College of North Wales, Bangor, Wales
• B.S.  Biological Science. University of East Anglia, Norwich, U.K.

Research Interests

Spatial distribution and range shifts of molluscs along the Atlantic due to climate change.  Development of shell ageing techniques and ageing of market species bivalves such as clams and scallops.  Physiological ecology of marine molluscs. Larval dispersal and settlement processes in estuarine and shallow water systems.

Academic Positions

2003-2012, Director for Research and Advisory Services, Virginia Institute of Marine Sciences
1987-present, Professor, William & Mary
1985-1987, Associate Professor, William & Mary
1981-1984, Associate Scientist, Woods Hole Oceanographic Institution (WHOI)
1977-1981, Assistant Scientist, WHOI
1975-1977, Post-Doctoral Fellow, WHOI
2022-present, Associate Graduate Faculty, University of Maine

Current Projects

• Population models of the Chesapeake oyster resources.
• Physiological and ecological studies of bivalve mollusc larvae in the field and laboratory.
• Dispersal processes in estuarine systems.
• Resource stock assessment by fishery dependent and fishery independent methods.
• Population assessment, life history and fishery management of offshore clam resources.
• Ecological restoration of oyster reef communities.
• Biology and ecology of veined rapa whelks in the Chesapeake Bay.
• Population demographics, growth rates, and age structure for Chesapeake hard clam resources.

Selected Fellowships and Grants (active in bold)

4/1/2024 – 3/31/29. Industry/University Cooperative Research Center: Science Center for Marine Fisheries (SCEMFIS) - Phase III. Eric N. Powell (University of Southern Mississippi) and Roger Mann (VIMS) $250.000. VIMS portion $125,000 - National Science Foundation plus $2,000,000 industry matching. Technical and travel support only from NSF portion.  Variable salary from industry portion.

10/1/2022 – 9/30/2023. The Virginia Seed Oyster Fishery: A Management Decision Tool. Roger Mann and Andrew Scheld co-PIs. Total $78,775. Virginia Marine resources Commission.  

6/1/2022-5/30/2024. Collaborative Research: Reconstructing bottom water temperatures from bivalves on the continental shelf: Holocene history as a window to the future in the Mid-Atlantic. Roger Mann (VIMS), Eric Powell (U. Southern Mississippi), Eileen Hofmann and John Klinck (Old Dominion Univ), Carolyn Ummenhoffer (WHOI). $396,469. VIMS portion $103,848. National Science Foundation 

5/1/2022-4/30/2023. Research Experience for a Undergraduate (REU) in the SCeMFiS projects.  – 3 students at $8,000 each. Total $24,000. National Science Foundation. Stipend, technical and travel support to REU participants.

9/1/2020-8/31/2023. Pew Foundation/LenFest Ocean Program. Support of a post doctoral investigator – climate change and fisheries. $210,418. Proposal generated by a 6 person team invited by Lenfest. Post doc situated at U. Wisconsin.

1/1/2021-7/1/2023. INTERN DCL NSF 18-102: Fisheries and climate change. NSF. $54,927. Stipend, technical and travel support to graduate student Alexis Hollander

11/1/2019-6/30/2023 (extended due to COVID). Research Experience for a Veteran (VET) in the SCeMFiS project entitled “Ocean quahogs (Arctica islandica) age structure.” - $10,000. National Science Foundation. Stipend, technical and travel support to veteran participant.

4/1/2019 – 3/31/24. Industry/University Cooperative Research Center: Science Center for Marine Fisheries (SCeMFiS). Eric N. Powell (University of Southern Mississippi) and Roger Mann (VIMS) VIMS portion - $500,000 from National Science Foundation plus $2,000,000 industry matching. Technical and travel support only from NSF portion.  Variable salary from industry portion.

3/1/2020-2/28/2023 (12 month no cost extension granted). Age based assessment in the sea scallop Placopecten magellanicus. Roger Mann and David Rudders, co-PI’s. $ 692,772. NOAA scallop RSA program

2/1/2021-6/30/2022. Research Experience for a Undergraduate (REU) in the SCeMFiS projects.  - $8,000. National Science Foundation. Stipend, technical and travel support to REUparticipant.

3/1/2018-2/28/2021 (12 month no cost extension).  The Effect of Density on the Growth, Yield and Reproduction of the Sea Scallop, Placopecten magellanicus. David Rudders and Roger Mann, co-PI’s. $320,279. NOAA scallop RSA program.

1/1/2018-3/312021. (no cost extension granted). Expanding Virginia’s oyster industry while minimizing user conflict.  Roger Mann, Marcia Berman, Melissa Southworth. PI. $261,288. DEQ - VA Coastal Zone Management Program.

10/1/2018-12/31/2018. Survey of clam resources in the HRBT expansion footprint. Roger Mann and Melissa Southworth co-PI’s. VA Department of Transportation.

11/1/2017-10/30/2018. Research Experience for an Undergraduate (REU) in the SCeMFiS project entitled “Ocean quahogs (Arctica islandica) age structure.” - $8,000. National Science Foundation. Stipend, technical and travel support to student participant.

9/1/2016-8/31/2019. Age structure and recruitment in the ocean quahog Arctica islandica. Roger Mann PI. $275,473 NOAA-SK.

3/1/2016-2/28/2019. Age structure and growth rate in the sea scallop Placopecten magellanicus. Roger Mann and David Rudders, co-PI’s. $608,626. NOAA scallop RSA program.

4/1/2013 – 3/31/2019 (includes one year no cost extension). Industry/University Cooperative Research Center: Science Center for Marine Fisheries (SCeMFiS). Eric N. Powell (University of Southern Mississippi) and Roger Mann (VIMS) VIMS portion - $300,000 from National Science Foundation plus $1,500,000 industry matching. Technical and travel support only from NSF portion.  Variable salary from industry portion.

3/1/2017-2/28/2018 in no cost extension. Shell/habitat dynamics in oyster restoration and fishery management. Roger Mann PI. $59,999. Chesapeake Bay Trust.

7/1/2014-6/30/2015. Participation of a high school teacher in the SCeMFiS project entitled “Ocean quahogs (Arctica islandica) recruitment and life history dynamics.” - $10,000. National Science Foundation. Salary, technical and travel support to teacher participant only. 

4/1/2013 – 3/31/2018. Industry/University Cooperative Research Center: Science Center for Marine Fisheries (SCeMFiS). Eric N. Powell (University of Southern Mississippi) and Roger Mann (VIMS) VIMS portion - $300,000. National Science Foundation plus $1,500,000 industry matching. Technical and travel support only from NSF portion.  Variable salary from industry portion.

10/1/2014-12/31/2014. Stock assessment of Virginia Oysters. Virginia Marine Resources Commission. Field personnel support only – reimbursable to $10,000.

7/1/2012-12/30/2014. Plumeri Award for Faculty Excellence. College of William and Mary. $10,000.

7/1/2013-9/30/2014. Shell budgets in the Virginia Chesapeake Bay: Quantitative estimates to support long term restoration and resource stabilization. Virginia Oyster Heritage Foundation. Roger Mann (VIMS) and James Wesson (VMRC) $30,000 (student and technical support).

10/1/2009-9/30/2012 (no cost extension through 9/30/2014). Collaborative Research: Climate Change and Responses in a Coupled Marine System. NSF. $613,707. Lead PI McCay (Rutgers) VIMS award $99,999. Mann 1.0 mo/yr. 

7/1/2011-6/30/2013 (no cost extension through 6/30/2014). A bay-wide approach to oyster stock assessment, estimates of vital rates and disease status. NOAA-NCBO. $ 336,193. Mann (lead PI) 1.0 mo/yr., with Ryan Carnegie (VIMS), James Wesson (VMRC), Mike Naylor and C. Dungan (MD DNR), Kennedy Paynter Jr. (UMD) and Howard Townsend (NOAA).

2/1/2012 – 1/31/2013. Planning Grant: Industry/University Cooperative Research Center for Mid-Atlantic Center for Fisheries Science (MaCFiS). Eric N. Powell (Rutgers) and Roger Mann (VIMS) $30,000. National Science Foundation. Technical and travel support only. 

1/1/2012-12/31/2013. Climate Change and the Fisheries Food Web in the Chesapeake Bay and coastal Atlantic Ocean. Robert Latour, Mary Fabrizio and Roger Mann. (VEE post doctoral support).

5/23/2011-12/21/2012. Shell substrate conditions and predator exclusion in oyster restoration. Virginia Oyster Heritage Foundation. Roger Mann (VIMS) and James Wesson (VMRC) $25,000 (technical support).

7/1/2011-6/30/2013 (no cost extension through 6/2014 to accommodate field work schedule). Oyster planting protocols to deter losses to cownose ray predation. NOAA-NCBO. $156,297. Mann (Lead PI) 1.0 mo/yr with James Wesson (VMRC), A.J. Erskine (Cowart Seafood), Tommy Leggett and Bill Goldsborough (CBF). 

11/1/2010-12/31/2013 – (final report in preparation, due 3/31/2014)  Shell budgets as a tool in oyster restoration and fishery management – application in Louisiana Primary State Seed Grounds. $150,000. National Fish & Wildlife Foundation. Mann (lead PI) 0.5 mo/yr. 

Selected Publications

• LeClaire A.M, E. N. Powell, R. Mann, K. M. Hemeon, S. M. Pace, V. Saba, H. du Pontavice, J. R. Sower. (2023). Temporal and Spatial Comparisons of Ocean Quahog (Arctica islandica) growth on the Mid Atlantic Continental Shelf: From the Neoglacial through the Twentieth Century. Continental Shelf Research. https://doi.org/10.1016/j.csr.2023.105143

• LeClaire, A.M., E.N. Powell, R. Mann, T. Redmond. (2023). Taphonomic indicators of dead ocean quahog (Arctica islandica) shell age in the death assemblage of the Mid-Atlantic Bight continental shelf. Palaios 38:305-324

• Stromp, S.L., E.N. Powell, R. Mann (2023).Evaluation of the Degree of Co-Occurrence of Atlantic Surfclams (Spisula solidissima) and Ocean Quahogs (Arctica islandica) in the Expanding Northwestern Atlantic Boreal/Temperate Ecotone: Implications for Their Fisheries. J. of Shellfish Research, 42(1):61-75 (2023). https://doi.org/10.2983/035.042.0107

• Stromp S.L., A. M. Scheld, J. M. Klinck, D. M. Munroe, E. N. Powell, R. Mann, S. Borsetti, E. E. Hofmann (2023). Interactive effects of climate change-induced range shifts and wind energy development on future economic conditions of the Atlantic surfclam fishery. Marine and Coastal Fisheries. https://doi.org/10.1002/mcf2.10232

• LeClaire, A.M., E.N. Powell, R. Mann, T. Redmond. (2023). Taphonomic indicators of dead ocean quahog (Arctica islandica) shell age in the death assemblage of the Mid-Atlantic Bight continental shelf. Palaios 38:305-324

• Sower J. R., E. N. Powell, K. M. Hemeon, R. Mann, S. M. Pace (2023). Ocean quahog (Arctica islandica) growth rate analyses of four populations from the Mid-Atlantic Bight and Georges Bank. Continental Shelf Research. https://doi.org/10.1016/j.csr.2023.105076

• Sower J. R., E. N. Powell, R. Mann, K. M. Hemeon, S. M. Pace, T. E. Redmond. (2023) Examination of spatial heterogeneity in population age frequency and recruitment in the ocean quahog (Arctica islandica Linnaeus 1767). Mar Biol 170, 38 (2023). https://doi.org/10.1007/s00227-023-04184-6

• Hemeon K.H., E. N. Powell, J. M. Klinck, R. Mann, S. M. Pace. (2023). Regional growth rates and growth synchronicity between two populations of Arctica islandica in the western Mid-Atlantic (US). Marine Biology. http://dx.doi.org/10.1016/j.ecss.2023.108412

• Bi R., R. Mann, K. E. Mills, V. Saba, J. Wiedenmann, O. P. Jensen (2023).  Empirical estimates of inter-assessment uncertainty for marine fish and invertebrate stocks. Fish and Fisheries. https://doi.org/10.1111/faf.12714

• Hemeon K.M., E. N. Powell, S. M. Pace, T. E. Redmond, R. Mann. (2023) Population dynamics of Arctica islandica off Long Island (USA): an analysis of sex-based demographics and regional comparisons. Marine Biology. https://doi.org/10.1007/s00227-023-04176-6

• LeClaire A.M , E. N. Powell, R. Mann, K. M. Hemeon, S. M. Pace, J. R. Sower, T. E. Redmond. (2022) Historical biogeographic range shifts and the influence of climate change during the Holocene on ocean quahogs (Arctica islandica) in the Mid-Atlantic Bight. The Holocene 10.1177/0959683622110127, 13 pp.

• Sower J. R., E. Robillard, E.N. Powell, K. M Hemeon, R. Mann (2022). Defining Patterns in Ocean Quahog (Arctica islandica) Sexual Dimorphism along the Mid Atlantic Bight. J. Shellfish Research 41(3):335-348. https://doi.org/10.2983/035.041.0304

• Hemeon K.H., E.N. Powell, S.M. Pace, T.E. Redmond, R. Mann. (2022). Population dynamics of Arctica islandica at Georges Bank (USA): an analysis of sex-based demographics. J. Marine Biological Association UK

• Mann R., E.N. Powell, D.M. Munroe. (2020). The Case of the “Missing” Arctic Bivalves and the Walrus, the Biggest [Ignored] Clam Fishery on the Planet. Journal of Shellfish Research. 39(3):1-9. https://doi.org/10.2983/035.039.0301 
• Powell E.N., R. Mann, M. C. Long, J. R. Timbs, K. M.  Kuykendall. (2020)^.  The conundrum of biont-free substrates on a high-energy continental shelf: Burial and scour on Nantucket Shoals, Great South Channel. Estuarine Coastal Shelf Science. https://doi.org/10.1016/j.ecss.2020.107089 
• Cronin K.E., S.E. Walker, R. Mann; A.S. Chute, M.C. Long; S.S. Bowser (2020). Growth and longevity of the Antarctic scallop Adamussium colbecki, an ecosystem engineer, under annual and multi-annual sea ice.  Antarctic Science. 32 (6) 466-475.  https://doi.org/10.1017/S0954102020000322] 
  
• Powell, E.N., J. Trumble, R. Mann, M.C. Long, S. Pace, J. Timbs, K. Kuykendall. (2020). Growth and Longevity in Surfclams East of Nantucket: Range Expansion in Response to the Post-2000 Warming of the North Atlantic. Continental Shelf Research. 195. 104059. 10.1016/j.csr.2020.104059. https://doi.org/10.1016/j.csr.2020.104059
  
• Powell, E.N., R. Mann, K. Kuykendall, M.C. Long, and J. Timbs. (2019). The intermingling of benthic macroinvertebrate communities during a period of shifting range: The “East of Nantucket” Atlantic Surfclam Survey and the existence of transient multiple stable states. Marine Ecology. 40. e12546. 10.1111/maec.12546.
• Lockwood, Rowan and Mann, Roger, "A conservation palaeobiological perspective on Chesapeake Bay oysters" (2019). VIMS Articles. 1827. https://scholarworks.wm.edu/vimsarticles/1827
• Timbs, J.R., E.N. Powell, and R. Mann. (2019). Changes in the spatial distribution and anatomy of a range shift for the Atlantic surfclam Spisula solidissima in the Mid-Atlantic Bight and on Georges Bank. Marine ecology. 620 77-97. doi:https://doi.org/10.3354/meps12964.
• Hennen, D., R. Mann, D. Munroe, E.N. Powell. (2018). Biological reference points for Atlantic surfclam ( Spisula solidissima) in warming seas. Fisheries Research. 207. 126-139. 10.1016/j.fishres.2018.06.013.
• Pace, S., E.N. Powell, R. Mann. (2018). Two-hundred year record of increasing growth rates for ocean quahogs (Arctica islandica) from the northwestern Atlantic Ocean. Journal of Experimental Marine Biology and Ecology. 503. 8-22. 10.1016/j.jembe.2018.01.010. 
• Timbs, J.R., E.N. Powell, and R. Mann. (2018). Assessment of the Relationship of Stock and Recruitment in the Atlantic Surfclam Spisula solidissima in the Northwestern Atlantic Ocean. VIMS Articles. 1338. 
• Hofmann, E.E., E.N. Powell, J.M. Klinck, D.M. Munroe, R. Mann, D.B. Haidvogel, D.A. NarvÁEz, X. Zhang, and K.M. Kuykendall. (2018). An overview of factors affecting distribution of the Atlantic surfclam (Spisula solidissima), a continental shelf biomass dominant, during a period of climate change. J. Shellfish Res. 37:821-831
• Pace, S. M., E.N. Powell, R. Mann, M.C. Long, and J.M. Klinck. (2017). Development of an Age-Frequency Distribution for Ocean Quahogs (Arctica islandica) on Georges Bank. J. Shellfish Res. 36: 41-53.
• Pace, S., E.N. Powell, R. Mann, M.C. Long. (2017). Comparison of age–frequency distributions for ocean quahogs Arctica islandica on the western Atlantic US continental shelf. Marine Ecology Progress Series. 585. 10.3354/meps12384.
• Powell, E.N., R. Mann, K.A. Ashton-Alcox, K.M. Kuykendall, and M.C. Long. (2017). “Can we estimate molluscan abundance and biomass on the continental shelf?” Estuarine, Coastal and Shelf Science. 198: 213-224
• Powell, E.N., J.M. Klinck, E.E. Hofmann, P. Moreno, K. Kuykendall, D.M. Munroe, and R. Mann (2016). Captains' response to a declining stock as anticipated in the surfclam (Spisula solidissima) fishery on the U.S. Mid-Atlantic coast by model evaluation, Ocean & Coastal Management, 134, pp. 52-68. https://doi.org/10.1016/j.ocecoaman.2016.09.012
• Munroe, D., D. Narvaez, D. Hennen; E. Hofmann, L. Jacobsen, R. Mann, E. Hofmann, E.N. Powell, J. Klinck (2016). The Roles of Fishing and Bottom Water Temperature as Drivers of Change in Maximum Shell Length in Atlantic Surfclams (Spisula solidissima) Estuarine Coastal and Shelf Sciences. 170: 112–122
• Powell, E.N., R. Mann, K. A, Ashton-Alcox, Y. Kim and D. Bushek (2015). The Allometry of Oysters: Spatial and Temporal Variation in the Length-Biomass Relationships for Crassostrea virginica.  J. Mar. Biol. Ass. UK
• Powell, E. N., J. M. Klinck, D. M. Munroe, E. E. Hofmann, P. Moreno, R. Mann. (2015).  The Value of Captains’ Behavioral Choices in the Success of the Surfclam (Spisula solidissima) Fishery on the U.S. Mid-Atlantic Coast: a Model Evaluation.
• Zhang, X., D. Haidvogel, D. M. Munroe, E. N. Powell, J. M. Klinck, R. Mann, F. Castruccio. (2015). Modeling Atlantic surfclams, Spisula solidissima, larval connectivity within the Middle Atlantic Bight: Model development, larval dispersal and metapopulation connectivity. Estuarine, Coastal and Shelf Science.
• Narváez. D. A., E. E. Hofmann, D. M. Munroe,  J. M. Klinck, E. N. Powell, R. Mann,  E. Curchitser. (2015). Long-term dynamics in Atlantic surfclam (Spisula Solidissima) populations: The role of bottom water temperature.  J. Mar. Sci.
• Kelsey M. Kuykendall, K.M., P. Moreno, E. N. Powell, T. M. Soniat, S. Colley, R. Mann, D. M. Munroe. (2015). The exposed surface area to volume ratio: Is shell more efficient than limestone in promoting oyster recruitment?  J. Shellfish Research. 34 (2): 217-225.
• Walles, B., R. Mann, T. Ysebaert, K. Troost, P.M. J. Herman and A. Smaal (2015). Demography of the ecosystem engineer Crassostrea gigas, related to vertical reef accretion and reef persistence.  Estuarine, Coastal and Shelf Science. 154: 224-233. https://doi.org/10.1016/j.ecss.2015.01.006.
• Soniat, T. N., N. Cooper, E. N. Powell, J. M. Klinck, M. Abdelguerfi, S. Tu, R. Mann and P. Banks. (2014). Estimating sustainable harvest of Eastern Oysters, Crassostrea virginica. J. Shellfish Research 33: 381-394. 
• Mann, R., M. Southworth, R. Carnegie and R. Crockett. (2014). Temporal variation in fecundity and spawning in the eastern oyster, Crassostrea virginica, in the Piankatank River, Virginia. J. Shellfish Research 33(1): 167-176.
• Mann, R., D.M. Munroe, E.N. Powell, E.E. Hofmann and J.M. Klinck. (2013). Bivalve molluscs: barometers of climate change in Arctic marine systems. In Responses of Arctic Marine Ecosystems to Climate Change. In:  F.J. Mueter, D.M.S. Dickson, H.P. Huntington, J.R. Irvine, E.A. Logerwell, S.A. MacLean, L.T. Quakenbush, and C. Rosa, Alaska Sea Grant, University of Alaska Fairbanks,  doi:10.4027/ramecc.2013.04.
• Harding, J. M., M. A. Unger, R. Mann, E. A. Jestel, and C. Kilduff. (2013). Rapana venosa as an indicator species for butyltin exposure over decadal and seasonal scales. Marine Biology. DOI 10.1007/s00227-013-2292-7
• Mann, R. (2013). Restoring nature’s coastal architects: a reality check. Ecological Restoration. 31(3) 323-324.
• Mann, R and M. W. Luckenbach. (2013).  Sperm swimming speeds in the oyster Crassostrea virginica (Gmelin 1791). J. Shellfish Research. 32(1): 1-4
• Waldbusser, G.G., E. N. Powell, and R. Mann. (2013). Ecosystem effects of shell aggregations and cycling in coastal waters: An example of Chesapeake Bay oyster reefs. Ecology 94(4): 895-903
• Munroe, D.M., E.N. Powell, R. Mann, J. M. Klinck, and E. E. Hofmann. (2013). Underestimation of primary productivity on continental shelves: evidence from maximum size of extant surfclam (Spisula solidissima) populations. Fisheries Oceanography. 22: 220–233. doi: 10.1111/fog.12016
• Harding, J.M.,  E.N. Powell, R. Mann and  M.J. Southworth^. (2013). Variations in Eastern oyster (Crassostrea virginica) sex ratios from three Virginia estuaries related to growth and mortality. Jour. Mar. Biol. Assoc. U.K. 93(2): 519-531.
• Harding, J. M., M.J. Southworth, Mann, R., and J. Wesson. (2012).  Comparison of Crassostrea virginica Gmelin (Eastern Oyster) recruitment on constructed reefs and adjacent natural Oyster bars over decadal time scales. Northeastern Naturalist. 19(4): 627-646. 
• Harding, J.M., W. M. Walton, C. M. Trappani, M. G. Frick and R. Mann. (2011). Sea turtles as potential dispersal vectors for non-indigenous species; The Veined Rapa Whelk as an Epibiont of Loggerhead Sea Turtles. Southeastern Naturalist. 10(2): 233-244
• Kennedy, V.S., D. L. Breitburg, M. C. Christman, M. W. Luckenbach, K. Paynter, J. Kramer, K. G. Sellner, J. Dew-Baxter, C. Keller, and R. Mann.^. (2011). Lessons Learned from Efforts to Restore Oyster Populations in Maryland and Virginia, 1990-2007. J. Shellfish Research. 30(3):719-732
• Harding, J. M., Mann, R., M. Southworth, and J. Wesson. (2010).  Management of the Piankatank River, Virginia, in support of oyster (Crassostrea virginica, Gmelin 1791) fishery repletion. J. Shellfish Research 29(4): 1-22
• Southworth, M.,  J. M. Harding, R. Mann and J. Wesson. (2010). Oyster (Crassostrea virginica Gmelin 1791) population dynamics on public reefs in the Great Wicomico River, Virginia, USA. J. Shellfish Research 29(2): 271-290.
• Harding, J. M., H. J. Spero, R. Mann, G. Herbert and J. Sliko. (2010).  Reconstructing early 17th century estuarine drought conditions from Jamestown oysters. Proc. National Academy Sciences. VIMS Articles. 952.
• Harding, J.M. and R. Mann. (2010). Distribution, size, and sex ratio of mature blue crabs Callinectes sapidus Rathbun 1896, from a Chesapeake Bay tributary in relation to habitat type and environmental factors. Bull. Mar Sci. 86(1): 75-91.
• Mann, R., J. M. Harding. and M. Southworth. (2009). Reconstructing pre-colonial oyster demographics in the Chesapeake Bay, USA. Estuarine, Coastal and Shelf Science 85: 217-222
• Harding, J. M., R. Mann, P. Moeller and M. S. Hsia. (2009). Mortality of the Veined Rapa Whelk, Rapana venosa, in relation to a Bloom of Alexandrium monilatum in the York River, USA. J. Shellfish Research. 28(2):
• Mann, R., M. Southworth, J.M. Harding and J. Wesson. (2009). Population studies of the native oyster Crassostrea virginica (Gmelin) in the James River, Virginia, USA. J. Shellfish Research. 28(2): 1-30
• Harding. J. M., R. Mann and M. Southworth. (2008). Shell length at age relationships in James River oysters (Crassostrea virginica) collected four centuries apart. J. Shellfish Research. 27(5): 1109-1115
• Harding, J. M., Gera, S. M., and R. Mann. (2008). Radula morphology in veined rapa whelks (Rapana venosa, Valenciennes 1846, Muriciade). The Nautilus 122(4):217-227
• Harding, J. M., R. Mann and C. Ware-Kilduff. (2008). Influence of environmental factors and female size on reproductive effort in a temperate marine gastropod Rapana venosa (Muricidae: Valenciennes 1846). Marine Biology 155(6): 571-581
• Harding, J. M., S. E. King. E. N. Powell, and R. Mann. (2008).  Decadal trends in age structure and recruitment patterns of ocean quahogs Arctica islandica from the Mid-Atlantic Bight in relation to water temperature. J. Shellfish Research 27(4): 667-690
• Varnell, L., D.M. Bilkovic, J.Olney, D.Evans, H.Wang, C.Hershner, R. Mann. (2008).  Estuarine surface water allocation: A case study on the interactive role of science in support of management.  Environmental Science and Policy 11(7): 602-612.
• Harding, J.M., Mann, R, Kingsley-Smith, P and Savini, D. (2007).  Comparison of predation signatures left by Atlantic oyster drills (Urosalpinx cinerea Say; Muricidae) and veined rapa whelks (Rapana venosa Valenciennes, Muricidae) in bivalve prey.  Journal of Experimental Marine Biology and Ecology.  352: 1-11
• Walter, J. F. III, M. C. Christman, J. M. Hoenig, and R. Mann. (2007).  Combining data from multiple years or areas to improve variogram estimation. Environmetrics 18: 1-16
• Mann, R. and E. N. Powell. (2007). Why oyster restoration goals in the Chesapeake Bay are not and probably cannot be achieved. J. Shellfish Research. 26(4): 905-917.
• Harding, J.M, Mann, R. and Ware-Kilduff, C.  (2007). The effects of female size on fecundity in a large marine gastropod Rapana venosa (Muricidae).  Journal of Shellfish Research. 26(1):33-42.
• Olney, J.E., D. M. Bilkovic, C. H. Hershner, L. M. Varnell, H. Wang and R. Mann. (2006). Six fish and 600,000 Thirsty Folks – A Fishing Moratorium on American Shad Thwarts a Controversial Municipal Reservoir Project in Virginia USA. American Fisheries Society Symposium, 2006: 587-598.
• Mann, R, Harding, JM, and E. Westcott. (2006).  Occurrence of imposex and seasonal patterns of gametogenesis in the invading veined rapa whelk Rapana venosa (Valenciennes, 1846) from Chesapeake Bay, USA. Marine Ecology Progress Series.  310: 129-138.
• Mann, R. (2006). The Local Costs To Ecological Services Associated With High Seas Global Transport. Pp 25-38 In: The Ecology of Transportation: Managing Mobility for the Environment J. Davenport and J. Davenport (eds). Kluwer Press.
• Harding, J. M. and R. Mann. (2006). Age and growth of Wild Suminoe (Crassostrea ariakensis, Fugita 1913) and Pacific (C. gigas, Thunberg 1793) Oysters from Laizhou Bay, China. J. Shellfish Res. 25(1): 73-82. 
• Harding, J. M. and R. Mann. (2005). Veined rapa whelk (Rapana venosa) range extensions in the Virginia waters of the Chesapeake Bay, USA. J. Shellfish Res. 381-386. 
• Powell, E.N. and R. Mann. (2005).  Evidence of recent recruitment in the ocean quahog Arctica islandica in the Mid-Atlantic Bight. J. Shellfish Res. 24(2): 517-530
• Mann, R., J. M. Harding, M. J. Southworth, J. A. Wesson. (2005)  Northern Quahog (Hard clam) Mercenaria mercenaria abundance and habitat use in Chesapeake Bay. J. Shellfish Res. 24(2): 509-516
• Mann, R., M. Southworth, J.M. Harding, and J. Wesson. (2004). A comparison of dredge and patent tongs for estimation of oyster populations. Journal of Shellfish Research. 23(2): 387-390.
• Southworth, M. and R. Mann. (2004). Decadal scale changes in seasonal patterns of oyster recruitment in the Virginia sub estuaries of the Chesapeake Bay. Journal of Shellfish Research. 23(2): 391-402.
• Mann, R. and D. Evans. (2004). Site selection for oyster habitat rehabilitation in the Virginia portion of the Chesapeake Bay: A commentary. Journal of Shellfish Research. 23(1): 41-49.
• Harding, J. M. and R. Mann. (2003). Influence of habitat on diet and distribution of striped bass (Morone saxatilis) in a temperate estuary. Bulletin of Marine Science. 72(3): 841-851.
• Mann, R. and J. M. Harding. (2003). Salinity tolerance of larval Rapana venosa: Implications for dispersal and establishment of an invading predatory gastropod on the North American Atlantic coast. Biological Bulletin. 204: 96-103.
• Baker, P.K. and R. Mann. (2003). Late stage bivalve larvae in a well-mixed estuary are not inert particles. Estuaries. 26(4A): 837-845.
• Bartol, I.K., R. Mann, and M. Vecchione. (2001). Distribution of the euryhaline squid Lolliguncula brevis in the Chesapeake Bay: Effects of selected abiotic factors. Marine Ecology Progress Series. In press.
• Harding, J.M. and R. Mann. (2001). Oyster reefs as fish habitat: Opportunistic use of restored reefs by transient fishes. Journal of Shellfish Research. 20(3): 951-959.
• Bartol, I.K, R. Mann, and M.R. Patterson. (2001). Aerobic respiratory costs of swimming in the negatively buoyant brief squid Lolliguncula brevis. Journal of Experimental Biology. 204:3639-3653.
• Mann, R. (2000). Restoring the oyster reef communities in Chesapeake Bay: a commentary. Journal of Shellfish Res. 19(1): 335-339.
• Mann, R. and J. M. Harding. (2000). Invasion of the North American Atlantic coast by a large predatory Asian mollusc. Biological Invasions. 2:7-22.
• Harding, J.M. and R. Mann. (1999). Observations on the biology of the veined rapa whelk, Rapana venosa (Valenciennes, 1846) in the Chesapeake Bay. Journal of Shellfish Research. 18(1): 9-17.

Honors, Prizes and Awards

• School of Marine Science Outstanding Faculty Award in Advisory Service, 2022

• At-large member of the Mid-Atlantic Fisheries Management Council representing Virginia.  Appointed by the Secretary of Commerce, First Term 2016-2019
• Governor's appointment to the Commonwealth's Aquaculture Advisory Board, Second Term 2013-2016 
• Plumeri Faculty Award, William and Mary 2012
• Chesapeake Bay Commission Tribute to Excellence, 1994, for “...official recognition of significant contributions to the management and protection of the Chesapeake Bay.” 
• School of Marine Science Outstanding Teacher Award, 2001
• Kirby Lang Visiting Professor in Marine Science, University College of North Wales, Bangor, Marine Science Laboratories, Menai Bridge, Wales, 2003
• Honored Life Member, National Shellfisheries Association, 2008
• Editor, Journal of Shellfish Research from 1982 -1986. Editorial board (current): Aquaculture (Elsevier), Journal of Shellfish Research

Offices in professional societies:

National Shellfisheries Association (in chronological order): 

Publications Committee member (1980-present), Publications Committee (2021 – present), Chair, Journal Editor (J. Shellfish Research) (1982-86), Editorial Board (1982-present), Vice President (1986-87), Program Chairman (1987 AGM), President Elect (1987-88), President (1988-89), AGM. Site and Program Organizer 1989-90, Constitutional Revision Committee 1990-91, Chair, Honorary Membership Committee 1991-92 and 1995-96.

Current and Graduated Students

Current Students:

Alexandria Marquardt. PhD candidate. Dynamics of oyster populations.

Kaitlyn Clark. PhD candidate, co-mentor with David Rudders. Density dependent growth in the scallop Placopecten magellanicus.

Graduated Students:

Alexis Hollander. 2022. M.S. Juvenile growth rates in the Atlantic surfclam Spisula solidissima over a multi-decadal year time frame in the Mid Atlantic Bight.

David Rudders 2010. Ph.D. (I mentored David during the final year of his work including the writing based on years of work after his major professor William DuPaul retired in 2009). Incorporating industry based dredge surveys into the assessment of the sea scallop, Placopecten magellanicus

Stefanie Gera. 2010. M.S. Egg Capsule Hatching Success in Rapana venosa and Urosalpinx  cinerea  in Relation to Temperature and Salinity.

Emily Chandler, 2007. M.S. Genetically monomorphic invasive populations of the rapa whelk, Rapana venosa.

Adriana Picariello 2006. M.S. The effects of climate change on the population biology of the surf clam, Spisula solidissima, in the Middle Atlantic Bight.

Elizabeth Darrow  (co-chair with Mark Luckenbach,). 2004 M.S. Ecological energetics of the hard clam.

Doolittle, Daniel. (co-chair with Mark Patterson). 2003. M.S. Development of a method for the identification and quantification of biological targets detected by sidescan sonar

Catherine Ware, 2002. M.S. Temporal and spatial variation in reproductive output of the veined rapa whelk (Rapana venosa) in the Chesapeake Bay.

Arminda Gensler. (co-chair with John Graves). 2001, M.S. Genetic Investigations of Interspecific and Intraspecific Relationships within the Genus Rapana.

Erica Westcott, 2001. M.S. Seasonal reproductive activity of the invading gastropod Rapana venosa in the Chesapeake Bay

Rebecca Green, 2001. M.S. Morphological variation of three populations of the Veined Rapa Whelk, Rapana venosa, an invasive predatory gastropod species.

Juliana M. Harding.  2000. Ph.D. Ecological interactions between benthic oyster reef fishes and oysters.

Ian Bartol. 1999. (co chair with Mark Patterson) Ph.D. Swimming Physiology of the Brief Squid, Lolliguncula brevis.

Melissa Southworth, 1998. M.A. Oyster Reef Broodstock Enhancement in the Great Wicomico River, Virginia

Sandra Brooke, 1996, M.A.  A comparison of natural and laboratory diets for the culture of marine invertebrate larvae: American Oyster, Queen Conch, and Milk Conch.

Elizabeth Keane - Shea, M.A. 1995 (co-chair with Mike Vecchione, Systematics Laboratory, National Museum, Smithsonian.)  The early life histories of three families of cephalopods, and an examination of the concept of a paralarvae. 

Ian Bartol, 1995, M.A.   Crassostrea virginica on constructed intertidal oyster reefs: Effects of tidal height and substrate level on settlement, growth and mortality. 

Patrick K. Baker, 1994, Ph.D.  Quantification of settlement and recruitment processes in bivalve molluscs. 

Shirley M. Baker, 1994, Ph.D.  Oyster (Crassostrea virginica) metamorphosis - Effects of low oxygen.

Robert A. Blaylock, 1992. Ph.D. Distribution, abundance and behavior of the Cownose Ray, Rhinoptera bonasus, in Lower Chesapeake Bay.

L. L. Castell, 1991, M.A.  Assessment of larval physiological condition using fluorescent, lipid specific stains.

G. Curtis Roegner, 1990, M.A.  Recruitment and growth of juvenile Crassostrea virginica (Gmelin) in relation to tidal zonation.

Kevin McCarthy, 1990, M.A.  The influence of swimming behavior of larval Crassostrea virginica (Gmelin).

Carrollyn Cox, 1988, M.A.  Seasonal changes in the fecundity of oysters Crassostrea virginica (Gmelin) from four oyster reefs in the James River, Virginia.

Bernardita Campos, 1988, M.A.  Swimming response of larvae of three mactrid bivalves to different salinity gradients.

David Eggleston, 1988, M.A.  Predator-prey dynamics between the blue crab, Callinectes sapidus Rathbun, and juvenile oysters, Crassostrea virginica (Gmelin).

Ellen Pafford, 1988, M.A.  Distribution and taxonomy of endolithic algae occurring in the shells of Crassostrea virginica in the lower James River, Virginia.

Courses Taught/Teaching

• Malacology, Larval Ecology, Biological Invasions