Virginia State Boundary
No legend
Oyster Reef Names
Oyster Reefs
No legend
Virginia River Basins
No legend
Intro Methods Help Summaries Literature

Introduction to VOSARA

The Virginia Oyster Stock Assessment and Replenishment Archive (VOSARA) data web site is designed to provide graphic summaries of the status of oyster stocks in the Virginia subestuaries of the Chesapeake Bay. Data originates in annual stock assessments performed by a collaboration between the Virginia Institute of Marine Science (VIMS) and the Virginia Marine Resources Commission (VMRC). Summaries provide data by year and historical reef boundaries within each of the described subestuaries. Oyster populations are described in units of density, oysters per unit area, for each of three size classes: spat are young of the year (≤ 35mm length), small are submarket size (> 35 mm and < 76mm length), and market size (≥ 76mm length) oysters.

For questions or comments about anything on this site please contact: Melissa Southworth (melsouth@vims.edu).

Preferred citation:
Southworth, M. and R. Mann. 2024. Virginia Oyster Stock Assessment and Replenishment Archive (VOSARA). Viriginia Institute or Marine Science, Gloucester Point VA https://cmap22.vims.edu/VOSARA/

Updated June 4, 2024

Oysters are collected during the fall (late September through December) from natural oyster reefs (public Baylor grounds) in the Virginia portion of the Chesapeake Bay. A quantitative sampling program is used at all reefs using a stratified random grid with documented oyster reefs (bars) forming the strata. In most cases, these reefs adhere to the locations and names used by Lieutenant J.B. Baylor, U.S. Coast and Geodetic Survey (Baylor, 1896) and subsequently resurveyed by Haven et al. (1981). Oysters are collected from the 43-ft long VMRC research vessel J.B. Baylor with a hydraulic patent tong. The open dimensions of the tongs are such that it samples one square meter of bottom. Upon retrieval of each sample (patent tong grab), oysters are counted and categorized as young of the year (hereafter spat; ≤ 35 mm), small (≥ 36 and ≤ 75 mm) or market (≥ 76 mm). In general this categorization is accurate, but the error associated with it may be exacerbated in areas where oysters regularly exhibit slow growth or in years of low salinity when the oysters may temporarily exhibit slow growth. Beginning in 2000 in the Rappahannock River, in 2002 for most other reefs that were sampled at that time and in 2003 for all reefs sampled, the shell material that was collected in each tong sample was categorized as either brown shell (shell that lies above the sediment water interface, including live oysters in all size-categories) or black shell (shell exhumed during the collection process) and the volume (L) in each category was recorded. The shell material measure (brown and black shell volumes) includes gravel or any other appropriate hard substrate that may be used for repletion/restoration on the various reefs. The procedures of Bros and Cowell (1987) were used to assure adequacy of sampling within each strata (reef). As of the most recent survey (fall 2023) there were a total of 219 individual reefs sampled as part of the survey, 197 of which are a part of VOSARA (reefs are added to VOSARA once five years of data have been collected).

Mean oyster density (number m-2) was calculated for each oyster reef by averaging the number of oysters in each category collected from all samples on a reef within a year. Mean shell volume was calculated for each oyster reef by averaging the volume of shell in each category collected from all samples on a reef within a year. For comparison purposes recruitment (young of the year or spat settlement) is categorized as: less than 50 spat m-2, low; 50 – 100 spat m-2, moderate; &ge100 spat m-2, good; > 400 spat m-2, very good. For each individual reef, years when particular reefs have received either a shell plant (SH: plant of clean shell on the bottom in the late spring or early summer), gravel plant (GR: plant of gravel of varying sizes on the bottom in the late spring or early summer), seed plant (SD: transplant of seed from another reef) or spat on shell plant (SOS) have been noted on the shell plots. Years when a particular reef had seed removed (SDR) as part of a project not associated with regular seed harvest by the industry is also noted on the shell plots. Seed is primarily small oysters and spat transplanted from another reef in the spring of a particular year, but may also include some larger oysters and blank shells. Spat on shell (SOS) is spat produced by allowing hatchery produced larvae to set on clean shell in a controlled tank environment and is typically planted in the summer.

Harvest information is included for each reef and was obtained from the Virginia Marine Resources Commission (VMRC) published regulations. The information is meant to inform the user on whether or not an area was opened or closed in a particular year and if opened what was allowed to be taken (seed or market oysters) as well as the type of gear used. The fact that an area was open for harvest in any particular year does not necessarily mean that oysters were harvested from that area, it simply means they legally could have been. The harvest year on public Baylor grounds in Virginia typically extends from October through April or May, so for harvest year 2024, the harvest extends from October of 2023 through May of 2024, but actual season lengths on different oyster reefs vary. The survey often overlaps with the harvest season in some years/rivers, so in any given year, data for a particular reef may have been collected pre-harvest, during the harvest or post-harvest. For the most recent regulations or for more information on individual reefs visit: https://mrc.virginia.gov/Regulations/FR720.shtm or https://webapps.mrc.virginia.gov/public/maps/chesapeakebay_map.php. Contact VMRC for additional information regarding past regulations.

  • Data is currently available for oyster reefs located in Great Wicomico River, James River, Lynnhaven Bay, Piankatank River, Pocomoke and Tangier Sounds, Rappahannock River, York River, and the main stem of the Chesapeake Bay.
  • Use the + and - buttons (left side of map) to zoom in and zoom out. Note: The zoom in and zoom out buttons are not available on a mobile device.
  • To return to the default map extent, click the home icon on the left side of the map.
  • Use the pull-down menu to view river choices; select river; river outline will highlight and map will zoom to river location. A mouse click anywhere on the map will remove the highlight.
  • In the upper right corner of the map, click the layer icon to open the layer list. Map layers can be checked on or off. River Basins will display at a scale of 1:750,000 and smaller. Oyster Reefs display at a scale of 1:750,000 and larger. Reef labels will display at 1:80,000 and larger. Reef labels show the VMRC name with the VIMS name in parentheses.
  • Legends for each layer are shown below each layer's name within the layer list. To hide or show a legend, click the icon to the right of the layer name.
  • Below the layer icon is the information icon . Click to show information about this viewer, methods used to collect the data, help on how to navigate the viewer, and summaries for the Chesapeake Bay Main Stem and each of the major tributaries containing oyster reef data.
  • On mobile devices, tap the map to dismiss/hide the information panel and the layer list panel.
  • Zoom into a location to show the oyster reefs. Click on an oyster reef containing data (those reefs without hatching) and an infowindow will pop-up in the lower left side of the map. The popup displays the reef name and acreage a brief summary about the reef, a harvest table, and graphs (oyster and shell). On mobile devices, the popup displays at the bottom of the screen. If the popup contents are not visible, tap the popup header to expand the contents.

CHESAPEAKE BAY MAIN STEM

There are seven public oyster reefs monitored in the Chesapeake Bay mainstem (approximately 100 acres) and one reef (Milford Haven; approximately 1 acre) that is located in the area known as “Hole in the Wall” (an area with Gwynn’s Island to the north and Mathew’s County to the south). Six of the seven reefs in the mainstem of the bay are part of the Deep Rock Management Area located east of Gwynn's Island. The remaining mainstem Chesapeake Bay reef (Blackberry Hang) and Milford Haven are both managed separately. Blackberry Hang is located just north of the mouth of the Great Wicomico River. There are between seven and twenty-two years of data included, depending on when monitoring began for a particular reef. Recruitment at all of these sites is typically very low, with only 27 out of the possible 121 year/site combinations having moderate (>50 spat m-2) or good (>100 spat m-2) recruitment. Since 2003, in the Deep Rock Management Area, 2015, 2019, 2021 and 2023 were notable year classes, on Blackberry Hang, 2015, 2016, 2019, 2022 and 2023 were notable and 2015 and 2022 were notable on Milford Haven. In general recruitment on all of the reefs in the mainstem of the bay has been higher since 2015. From 2003 to 2014, average recruitment on the Deep Rock Management Area reefs and Blackberry Hang ranged from 1 to 14 and from less than 1 to 21 spat m-2, respectively. In contrast, average recruitment from 2015 to 2023 in the two areas (Deep Rock Management Area and Blackberry Hang), ranged from 10 to 92 and from 15 to 146 spat m-2, respectively.

In recent years, the density of market oysters on all of these reefs has tended to be higher than in earlier years. From 2003 to 2008, the overall average density of market oysters on the reefs in the Deep Rock Management Area was less than 1 m-2, whereas from 2009 to 2023, the average density of market oysters ranged from 2 to 29 m-2 and has remained above 5 m since 2012 and above 10 m-2 since 2020. From 2004 to 2016 market oyster density on Blackberry Hang was never more than 5 m-2, but from 2017 to 2023, market oyster density ranged from 11 to 20 m-2.

GREAT WICOMICO RIVER

There are twenty-four public oyster reefs (approximately 193 acres) monitored in the Great Wicomico River. Seventeen of these reefs are located upriver of Sandy Point in the traditional seed harvest area with the other seven located downriver of Sandy Point in the harvest (market oysters) area. This area has been harvested with hand scrapes since 2006. There are between ten and twenty-four years of data included, depending on when monitoring began for a particular reef. Recruitment in the system was highly variable (ranging from a low of 0 spat m-2 to a high of 1051 spat m-2), both within the system at different sites in a given year and between years. Overall, there were five strong year classes observed during the monitoring period; 2002, 2006, 2012, 2016 and 2022 with an overall average of 101 (seven reefs), 167 (eighteen reefs), 189 (twenty-two reefs), 154 (twenty-four reefs) spat m-2, and 127 (twenty-four reefs), respectively. In general, there have been more small and market oysters in the system since 2007. In every year from 2000 to 2006, the average over all of the sites was less than 1 market oysters m-2 with less than 20 small and market oysters m-2 combined. From 2007 to 2023, the average over all of the sites in a given year ranged between 2 and 12 market oysters m-2 with 18 to 97 small and market oysters m-2 combined. The average market oysters over all twenty-four reefs has remained above 10 m-2 for the past three years (since 2021).

JAMES RIVER

There are thirty-four public oyster reefs (approximately 6,820 acres) monitored in the James River. Twenty-three of these are located upriver in the traditional seed, hand tong area, with the remaining eleven located downriver in the hand-scrape harvest (market oysters) area. One reef, Wreck Shoal, located in the seed area was designated as a sanctuary reef in 2010. There are between nine and twenty-six years of data included, depending on when monitoring began for a particular reef. Recruitment in the system over the monitoring period was highly variable (ranging from a low of 0 spat m-2 to a high of 1,319 spat m-2), both within the system at different sites in a given year and between years.

Recruitment on the twenty-three reefs located in the traditional seed area, has been especially good since 2008. From 1998 to 2007, there was only one year (2002) when the average density of spat over all twenty-three reefs was greater than 100 spat m-2. Since 2008, the average density of spat over all twenty-three reefs has been greater than 100 spat m-2 in fourteen out of the sixteen years. Only over the past two years (2021 and 2022) has this increased recruitment resulted in an increase in the overall number of markest oysters on these reefs. The average market oyster density over all of the reefs was at a twenty-six year high in 2023 (average of 24.3 m-2).

At the eleven sites located in the hand-scrape harvest area, the three strongest year classes were 2015, 2018, and 2019. Recruitment in this area of the river has generally been higher in recent years. From 2006 to 2014, the average recruitment over all of the sites ranged from 1.8 to 17.7 spat m-2, whereas from 2015 to 2023, the average recruitment over all the sites ranged from 50.0 to 252.7 spat m-2. In general, the density of small and market oysters combined has been higher since 2015 as well. From 2006 to 2014, the average small and market oysters combined over all of the sites ranged from 3.2 to 9.8 m-2, whereas from 2015 to 2023, the average ranged from 19.8 to 130.0 m-2. It should be noted that White Shoal was added in 2015 so from 2006 to 2014, there were only ten sites included in the calculations, versus eleven sites from 2015 to 2023. The overall average market oyster density on these eleven reefs has remained above 10 m-2 for the past three years. Also as noted in the specific text regarding Nansemond Ridge, this reef has been the focus of regular shell plants for the purpose of seed movement to other reefs within the Commonwealth since 2017.

LYNNHAVEN BAY

There are twelve public oyster reefs (approximately 48 acres) monitored in the Lynnhaven River system. All of these are designated as sanctuary reefs, closed to harvest. There are between eleven and fourteen years of data included, depending on when the particular reef was built and monitoring began. Eastern Branch 3 and Eastern Branch 4 were not sampled in 2017 and haven`t been sampled since 2018. Eastern Branch 1 and Eastern Branch 2 have not been sampled since 2016. The remaining reefs in the system were not sampled in 2019, 2021 and 2023. Recruitment was typically low (less than 50 spat m-2) throughout the system, but varied depending on location, with the Broad Bay area typically having the highest recruitment. The four exceptions to low recruitment occurred on Broad Bay 2 in 2016 (183 spat m-2) and 2022 (106 spat m-2) and on Linkhorn Bay 3 (101 spat m-2) and Linkhorn Bay 4 (80 spat m-2) in 2017. The strongest year classes in each of the different areas of the Lynnhaven River system were 2008 (Eastern Branch reefs), 2016 (Broad Bay reefs) and 2017 (Linkhorn Bay reefs). In Broad Bay, 2022 was also a notable year for recruitment. The reefs in the Eastern Branch are in a fairly dynamic area and the majority of the shells that were initially planted have essentially been buried by sand. From 2013 to 2015, greater than 31% of the oysters in the system were market size, with an overall system average of 12 to 15 market oysters m-2. This has varied since, with a mean over all the reefs in 2018, 2020 and 2022 of 13.3, 30.0 and 21.1 m-2 respectively. Market oysters accounted for 23.7% of the oyster population in 2018, 39.3% in 2020 and 23.1% in 2022.

PIANKATANK RIVER

There are eighteen public oyster reefs (approximately 227 acres) monitored in the Piankatank River. Five of these are seed reefs (most recent harvest on any of these reefs was in 2018), but the rest have either never been harvested or for the older reefs have not been harvested for at least twenty years or more. There are between nine and twenty-six years of data included, depending on when monitoring began for a particular reef. Recruitment in the system was variable (ranging from a low of 0 spat m-2 to a high of 308 spat m-2), both within the system at different sites in a given year and between years. Overall, there were three strong year classes observed in the system over the monitoring period. These were 1999, 2002, and 2010, with an average of 115 (eight reefs), 114 (nine reefs) and 110 (ten reefs) spat m-2 respectively. The most recent notable year was 2019 with 72 spat m-2. Following a fairly large die-off of oysters in all size classes, the oyster population in the Piankatank River was at very low levels in 2004 and 2005. The population of small and market oysters had started to rebound by 2007 (small oysters) and 2008 (market oysters) and in general have remained at higher levels since then. From 1998 to 2006, the overall average density of small oysters in the river ranged between 1.0 and 18 m-2 whereas from 2007 to 2023 the overall average ranged between 18 and 55 m-2. For market oysters, the overall average density in the river from 1998 to 2007 was consistently less than 1 m-2, while from 2008 to 2023 the density ranged between 2 and 14 market oysters m-2. The highest density of small oysters observed over the twenty-six year period occurred in 2021 and the highest density of market oysters observed occurred in 2022.

RAPPAHANNOCK RIVER

There are fifty-nine public oyster reefs (approximately 710 acres) monitored in the Rappahannock River. Seventeen of these are designated as sanctuary reefs with the remaining fourty-two located in the rotational harvest areas (Rappahannock River Rotation Areas 1-6; thirty-two reefs), in Rappahannock River Area 7 (eight reefs), Rappahannock River Area 8 (one reef) or the Corrotoman River Hand Tong Area (one reef). There are between six and twenty-four years of data included, depending on when monitoring began for a particular reef. Recruitment in the system was highly variable (ranging from a low of 0 spat m-2 to a high of 559 spat m-2), both within the system at different sites in a given year and between years. There was a seven-year period (2003 to 2009) when recruitment was consistently low (overall average across the river of less than 10 spat m-2), but recruitment over the past fourteen years (2010 to 2023) has improved, such that seven of the years had an average greater than 20 spat m-2 (six out the past nine years). In most years, recruitment tended to be slightly higher at the more downriver sites but in 2022 and 2023 recruitment was highest at the more mid to upriver sites. Average recruitment over the past two years at most of these sites was anywhere from 4 to 50 times higher than what is typical in that part of the river. From 2002 through 2021, the overall mean recruitment at the more upriver reefs in the Rappahannock River system (the reefs located upriver of Towle's Point) was never more than 15 spat m-2. The overall mean at these reefs in 2022 and 2023 was 108 and 50 spat m-2, respectively. Throughout the Rappahannock River system, three years, 2015, 2019 and 2022, had an overall mean recruitment (mean across all reefs monitored in a given year) greater than 50 spat m-2; 2023 was also notable with an overall average of 49 spat m-2. In general, the density of market oysters has been higher since around 2008. From 2000 to 2007, the overall density of market oysters ranged from less than 1 m-2 to 2 m-2. From 2008 to 2023, the overall density of market oysters in any given year ranged between 2 and 19 m-2, and has remained above 5 m-2 since 2012 and above 15 m-2 for the past three years.

TANGIER AND POCOMOKE SOUNDS

There are fourteen public oyster reefs monitored in Pocomoke Sound (approximately 472 acres) and thirteen monitored in Tangier Sound (approximately 166 acres). Nine of the reefs in Pocomoke Sound and twelve of the reefs in Tangier Sound are part of the Pocomoke and Tangier Sound Rotational Harvest areas. The remaining five reefs in Pocomoke Sound are located near the mouth of the Pocomoke River and are managed separately. The remaining reef in Tangier Sound (Cod Harbour) is a designated sanctuary reef. There are between nine and twenty-one years of data included, depending on when monitoring began for a particular reef. Recruitment in these systems is variable, both within the systems at different sites in a given year and between years. Recruitment was typically low or moderate, with two reefs in Tangier Sound having good (>100 spat m-2) recruitment in 2020 one having good recruitment in 2022 and two having good rectuitment in 2023. Recruitment over the monitoring period ranged from 0 to 91 spat m-2 in Pocomoke Sound and from less than 1 to 299 spat m-2 in Tangier Sound.

The strongest year class in Pocomoke Sound was 2019 (overall average was almost twice as high as the next highest year), but 2006, 2007, 2010, 2018, 2021 and 2023 were also notable years; these were all years with an overall average >15 spat m-2. In general, the density of small and market oysters increased in years following these recruitment events. From 2012 to 2017, recruitment was extremely low in this system with the overall density in all size classes reaching a low of 1.6 m-2 in 2015. Both small and market oyster densities have increased since, with around 13 and 8 small and market oysters m-2 in 2023, respectively.

The strongest year class in Tangier Sound was 2020. There were six years that had an overall average between 20 and 30 spat m-2 (2002, 2006, 2015, 2018, 2021 and 2022) and four that had an overall average greater than 35 spat m-2 (2007, 2019, 2020 and 2023). The overall average spat observed in Tangier Sound has been greater than 25 m-2 for the past six years (2018 through 2023). In the sixteen years prior, 2007 was the only year that had greater than 25 spat m-2. In general, the density of small and market oysters increased in years following each of the notable recruitment events, with the overall average of small oysters over the twenty-two years ranging between 2 and 37 m-2 and the overall average of market oysters ranging between less than 1 and 21 m-2. Average oyster market density has remained above 17 m-2 for the past three years.

YORK RIVER AND MOBJACK BAY

There are seven public oyster reefs monitored in the York River (approximately 257 acres) and five monitored in Mobjack Bay (approximately 129 acres). Three of the five reefs in the York River and three of the reefs in Mobjack Bay are part of the York River and Mobjack Bay rotational harvest areas. The remaining four reefs in the York River and the two reefs in the Mobjack Bay are designated as sanctuary reefs. There are between five and eighteen years of data included, depending on when monitoring began for a particular reef. Recruitment in these systems is variable, both within the systems at different sites in a given year and between years. Recruitment was typically low to moderate and ranged from less than 1 to 414 spat m-2 in the York River and from 0 to 744 spat m-2 in Mobjack Bay. During the monitoring period, 2019 was the strongest year class in the York River, with 2015 also being a notable year. The strongest year class in Mobjack Bay was 2019, with 2015 also being notable. Recruitment in 2019 in Mobjack Bay was two to four times higher than 2015 recruitment (the next highest year at all of the sites) and twelve to twenty-six times higher than the third highest recruitment year (recruitment varied between sites).

In the York River, the density of market oysters on the four sanctuary reefs trended upward over the fifteen-year monitoring period, with the average over the two reefs remaining above 11 m-2 since 2020. On the harvest reefs, the density of market oysters has been higher since 2012, when compared with earlier years; from 2009 to 2011, there were less than 1 market oysters m-2 in any given year, whereas the density from 2012 to 2023 ranged between 3 (2012 and 2015) and 13 (2022) market oysters m-2. The density of small and market oysters combined on all seven reefs in the York River was around 4 m-2 in 2009 and 2010, but began to increase in 2011 and has ranged between 22 and 33 m-2 for the past eight years (2016 to 2023).

In Mobjack Bay, the density of market oysters on all of the reefs varied a good bit over the monitoring period, ranging from an average low of less than 1 m-2 to a high of 11 m-2. On Brown's Bay #2, (added to the VOSARA site in 2018) market oyster density increased from less than 1 market oyster m-2 in 2013 through 2015, the first three years of sampling on the reef, to an average of 11 to 25 market oysters m-2 from 2016 to 2023. On Brown's Bay #1 (added to the VOSARA site in 2020) the density of market oysters has remained above 18 m-2 since 2021.

Suggested Literature

Baylor, J. B. 1896. Method of defining and locating natural oyster beds, rocks and shoals. Oyster records (pamphlets, one for each Tidewater, Virginia, County that listed the precise boundaries of the Baylor Survey). Board of Fisheries of Virginia. 770 pp.

Bros, W. E. & B. C. Cowell. 1987. A technique for optimizing sample size (replication). J. Exp. Marl. Biol. Ecol. 114:63-71.

Haven, D. S., J. P. Whitcomb, & P. Kendall. 1981. The present and potential productivity of the Baylor Grounds in Virginia. Virginia Institute of Marine Science, Special report Applied Marine Science and Ocean Engineering no. 243. Gloucester Point, VA: Virginia Institute of Marine Science. 154 pp.

Harding, J. M. & R. Mann. 2010. Observations of distribution, size, and sex ratio of mature blue crabs Callinectes sapidus Rathbun 1896 from a Chesapeake Bay tributary in relation to oyster habitat and environmental factors. Bull. Mar. Sci. 86(1):75-91.

Harding, J. M., R. Mann, & M. Southworth. 2008. Shell length-at-age relationships in James River, Virginia oysters (Crassostrea virginica) collected four centuries apart. J. Shellfish Res. 27(5):1109-1116.

Harding, J. M., E. N. Powell, R. Mann, & M. Southworth. 2013. Variations in eastern oyster (Crassostrea virginica) sex-ratios from three Virginia estuaries: protandry, growth and demographics. J. Mar. Biol. Assoc. U.K. 93(2):519-531.

Harding, J. M., M. Southworth, & R. Mann. 2010. Observations of blue crabs (Callinectes sapidus, Rathbun, 1896) on Shell Bar oyster reef, Great Wicomico River, Virginia, USA. J. Shellfish Res. Vol. 29, No. 4, 995-1004.

Harding, J. M., R. Mann, M. Southworth, & J. A. Wesson. 2010. Management of the Piankatank River, Virginia in support of oyster (Crassostrea virginica, Gmelin 1792) fishery repletion. J. Shellfish Res. 29(4):1-22.

Harding, J. M., M. Southworth, R. Mann, & J. A. Wesson. 2012. Comparison of Crassostrea virginica Gmelin (Eastern Oyster) recruitment on constructed reefs and adjacent natural oyster bars over decadal time scales. Northeast. Nat. 19(4):627-646

Mann, R. & D. Evans 1998. Estimation of oyster, Crassostrea virginica, standing stock, larval production, and advective loss in relation to observed recruitment in the James River, Virginia. J. Shellfish Res. 17:239-253.

Mann, R., J. M. Harding, & M. Southworth. 2009. Reconstructing pre-colonial oyster demographics in Chesapeake Bay, USA. Estuarine, Coastal and Shelf Science. 85(2009):217-222.

Mann, R., M. Southworth, R. B. Carnegie, & R. K. Crockett. 2014. Temporal variation in fecundity and spawning in the Eastern Oyster, Crassostrea virginica, in the Piankatank River, Virginia. J. Shellfish Res. 33(1):167-176.

Mann, R., M. Southworth, J. M. Harding, & J. A. Wesson. 2004. A comparison of dredge and patent tongs for estimation of oyster populations. J. Shellfish Res. 23(2):387-390.

Mann R., M. Southworth, J. M. Harding, & J. A. Wesson. 2009. Population studies of the native oyster, Crassostrea virginica, (Gmelin, 1791) in the James River, Virginia, USA. J. Shellfish Res. 28(2):193-220.

Mann, R., M. Southworth, J. Wesson, J. Thomas, M. Tarnowski, & M. Homer. 2021. Oyster shell production and loss in the Chesapeake Bay. J. Shellfish Res. 40(3):451-469.

Southworth, M., J. M. Harding, R. Mann and J. Wesson. 2010. Oyster (Crassostrea virginica Gmelin 1791) population dynamics on public reefs in the Great Wicomico River, Virginia, USA. J. Shellfish Res. 29(2):271-290.

Southworth, M. and R. Mann. 2004. Decadal scale changes in seasonal patterns of oyster recruitment in the Virginia subestuaries of the Chesapeake Bay J. Shellfish Res. 23(2):391-402.

20 km
20 mi
Earthstar Geographics
Powered by Esri